Areas of research interest and publications have included erythrocyte metabolism of domestic animals, Heinz body hemolytic anemias, serum haptoglobin alterations in disease states, iron deficiency anemia, methemoglobinemias, myeloproliferative and lymphoproliferative disorders, developmental hematology and iron metabolism in foals, hematology and iron metabolism of pregnant mares, and infectious agents of blood cells. Infectious agents studied include Mycoplasma haemofelis, Cytauxzoon felis, Anaplasma platys, Ehrlichia canis, Anaplasma phagocytophilum, Babesia canis and Leishmania donovani.
A longstanding area of interest is inherited defects of erythrocyte metabolism in domestic animals. Publications have resulted from in depth studies of clinical cases of pyruvate kinase deficiency in dogs, methemoglobin reductase deficiency in dogs and cats, glucose-6-phosphate dehydrogenase deficiency in horses, and flavin adenine dinucleotide deficiency in horses. Extensive experimental studies have been conducted on erythrocytes and muscles from adult and neonatal phosphofructokinase-deficient dogs. A new area of study focuses on the clinical chemistry and hematology of Florida manatees.